- Ecotoxicology ot terrestrial invertebrates
- Evolution of resistance to toxicants
- Biodiversity in agricultural landscapes
- Diversity and activity of microorganisms in reclaimed soils
- Effect of pollution, natural stressors and their interactions on soil microbial communities
- Shifts it tolerance of soil microbial communities in response to stressors
- Effect of toxicants on gene expression and enzyme activity in bank voles
- Toxicokinetics of trace metals in bank vole tissues
- Nature conservation