- Ecotoxicology of invertebrates (effect of toxic chemicals on life history, populations and communities; interactions among toxicants and natural stressors; evolution of resistance)
- Toxicology and ecotoxicology of small vertebrates (histopathology, biomarkers, genetic effects)
- Ecosystem ecotoxicology (effect of pollutants on litter decomposition and nutrient turnover)
- Terrestrial ecosystems ecology (organic matter decay, biogeochemistry, biodiversity)