- Life histories and life strategies of plants and animals: experimental, comparative, theoretical approach
- Thermal biology of ectotherms: performance, thermal limits, life strategies, oxygen limitations
- Adaptive evolution of cell size
- Evolutionary ecology of multitrophic interactions: mutualistic symbiosis, host-parasite, predator-prey, herbivore-plant
- Optimal resource allocation: the origin of diversity in body size, offspring size, growth patterns, ageing, reproductive modes