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  • Ecotoxicology of invertebrates (effect of toxic chemicals on life history, populations and communities; interactions among toxicants and natural stressors; evolution of resistance)
  • Toxicology and ecotoxicology of small vertebrates (histopathology, biomarkers, genetic effects)
  • Ecosystem ecotoxicology (effect of pollutants on litter decomposition and nutrient turnover)
  • Terrestrial ecosystems ecology (organic matter decay, biogeochemistry, biodiversity)